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Escaping conflict on Earth, an idealistic group of settlers arrive on a distant planet - Pax - with plans for a perfect society. The world they discover is rich with life, but this is not the Eden they were hoping for. The plants on Pax are smart - smart enough to domesticate, and even slaughter, its many extraordinary animals. To survive, the colonists realise that they must strike bargains of their own. But if they are to make Pax their home, they must go further, searching for a way to communicate and coexist with these utterly alien intelligences.

Over two hundred years after the first colonists landed on Pax, a new set of explorers arrives from Earth on what they claim is a temporary scientific mission. But the Earthlings misunderstand the nature of the Pax settlement and its real leader. Even as Stevland attempts to protect his human tools, a more insidious enemy than the Earthlings makes itself known. Stevland is not the apex species on Pax.